火箭浣熊（布莱德利·库珀 Bradley Cooper 饰）偷走了大祭司阿耶莎（伊丽莎白·德比齐 Elizabeth Debicki 饰）的能量电池，包括星爵（克里斯·帕拉特 Chris Pratt 饰）、卡魔拉（佐伊·索尔达娜 Zoe Saldana 饰）、德拉克斯（戴夫·巴蒂斯塔 Dave Bautista 饰）在内的一行人遭到了后者派出的舰队的袭击。一个神秘人物乘坐着飞船救下了银河护卫队的众人，而驾驶着飞船的不是别人，竟然正是星爵的亲生父亲伊戈（库尔特·拉塞尔 Kurt Russell 饰）。 在伊戈的带领下，星爵、卡魔拉和德拉克斯来到了由伊戈一手制造的星球上，而剩下的成员们则着手修复损坏的飞船。那边厢，阿耶莎委托勇度（迈克尔·鲁克 Michael Rooker 饰）抓住星爵替自己报仇，而卡魔拉亦渐渐发现了伊戈和善面目下隐藏的巨大阴谋。
In Italy, old, dirty, congested prisons full of violence and abuse are a dim and distant memory. Now prisoners serve their sentences in a state of deep sleep which renders them inoffensive and has drastically reduced reoffending rates. Hypersleep is an efficient, economically viable and reliable system. That is, until the day when David Damiani (Stefano Accorsi), a psychologist charged with monitoring prisoners’ mental states, finds himself faced with a prisoner about whom all information has been lost. It’s an unexpected event which triggers a chain of unforeseeable event, forcing David to face up to the ghosts of his very own past.
Four men working at a remote scientific outpost unravel the disturbing secrets about the facility's history, signaling the approach of a mysterious menace that will not only consume these men, but, perhaps, everyone on the planet.
In a dystopian future where all of humanity's memories have become government-regulated transactional products, mankind's crown-jewel memory is stolen - the answer to where we come from.
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